Leonard Hayflick, Ph.D.
Professor of Anatomy (Adj.)
University of California, San Francisco School of Medicine
36991 Greencroft Close
P. O. Box 89
The Sea Ranch, California 95497
Date and Place of Birth May 20, 1928, Philadelphia, Pennsylvania
Marital Status Married, 5 children
Address University of California, San Francisco
School of Medicine
Department of Anatomy
36991 Greencroft Close
P.O. Box 89
The Sea Ranch, California 95497
Telephone (707) 785-3181
FAX (707) 785-3809
John Bartram High School, Philadelphia, Pennsylvania, 1946
University of Pennsylvania, Phila, PA B.A. Microbiology and Chemistry, 1951
University of Pennsylvania, Phila, PA M.S. Medical Microbiology, 1953
University of Pennsylvania, Phila, PA Ph.D. Medical Microbiology and Chemistry, 1956
U.S. Army, Ordnance Department, Staff Sergeant, 1946-1948
Winner of competitive scholarship to Temple University, Philadelphia, Pennsylvania upon
completion of high school, February, 1946. Scholarship declined in order to attend
University of Pennsylvania, Department of Microbiology. University of Pennsylvania
Scholarship, September 1953 - June 1954.
University of Pennsylvania Scholarship, September 1954 - June 1955.
George Leib Harrison Fellowship in Medical Microbiology, September 1955 - June 1956.
James W. McLaughlin Research Fellowship in Infection and Immunity, Post-doctoral,
University of Texas Medical Branch, Galveston, Texas, August 1956 - March 1958.
Career Development Awardee, National Cancer Institute, National Institutes of Health,
Professor of Anatomy (Adj.) Research Cell Biologist
University of California, San Francisco Dept. of Veteran's Affairs, DVA Medical
Center,151E School of Medicine and Cell Biology and Aging Section
Department of Anatomy 4150 Clement St.
Parnassus Street San Francisco California , 94121 San Francisco, CA 94143-0452
Previous Positions Held
1. Director, Center for Gerontological Studies, College of Liberal Arts and Sciences,
University of Florida, 3357 Turlington Hall, Gainesville, Florida 32611 1981-1986
2. Professor of Zoology, Department of Zoology, School of Liberal Arts and Sciences,
University of Florida, Gainesville, Florida 32611 1981-1986
3. Professor of Microbiology and Immunology, Department of Microbiology and Immunology
College of Medicine, University of Florida, Gainesville, Florida 32611 1981-1986
4. Senior Research Cell Biologist, Bruce Lyon Memorial Research Laboratory, Children's
Hospital Medical Center, Oakland, California, 1976-81.
5. Professor of Medical Microbiology, Department of Medical Microbiology, Stanford
University School of Medicine, Stanford, California, 1968-76.
6. Assistant Professor of Research Medicine, Department of Research Medicine, University
of Pennsylvania School of Medicine, Philadelphia, PA, 1966-68.
7. Associate Member, The Wistar Institute of Anatomy and Biology, 36th and Spruce Streets,
Philadelphia, PA, 1958-68.
8. James W. McLaughlin Research Fellow in Infection and Immunity, Department of
Microbiology, University of Texas Medical Branch, Galveston, Texas. August, 1956 - April,
9. Research Assistant in Bacteriology, Sharp and Dohme, Division of Merck and Company,
Inc., June 1951 - June 1952.
1. Non-Resident Fellow, Institute for Higher Studies, Santa Barbara, California, 1973 -
2. Member, Editorial Board, "Executive Health Report", 1970 - Present.
3. Member, Editorial Board, "Mechanisms of Ageing and Development", 1972 -1998.
22. Member, Program Committee and Awards Committee, Gerontological Society of America,
23. Member, Virology and Cell Biology Study Section, American Cancer Society, 1974-1976.
24. Member, Subcommittee on the Mycoplasmataceae of the International Committee on
Bacteriological Nomenclature, 1965-1978.
25. Member, Advisory Committee, Bergey's Manual of Determinative Bacteriology, 1965-1978.
26. Program Committee, Western Gerontological Society, 1977-1979.
27. Member, Committee on International Relations, Gerontological
Society of America, 1980-1982.
28. Member, Public Policy Committee, Gerontological Society of America, 1980-1982.
29. Observer, White House Conference on Aging, 1981, Representing Gerontological Society
of America and State of California.
30. Member, Board of Director, Western Gerontological Society, 1981- 1983.
31. Offered First Directorship of National Institute on Aging, NIH, Bethesda, MD, April
1975. Offer declined.
32. President, Gerontological Society of America, 1982-1983.
33. Director, Center for Gerontological Studies, College of Liberal Arts and Sciences,
University of Florida, 1981-1985.
34. Member, Advisory Board, International Exchange Center for Gerontology, State of
Florida University System, University of South Florida, Tampa, 1982-1986.
35. Secretary, Cell Culture Committee, International Association of Biological
36. Member, Research Advisory Committee, Teachers Insurance and Annuity Association of
America - College Retirement Equities Funds, New York, N.Y., 1974 - 1980.
37. Elected Councillor, Society for Experimental Biology and Medicine, 1984-1988.
38. Member of Sandoz Jury for Sandoz Prize in Gerontology and Geriatrics, 1985 - 1989.
39. Member of Council of the International Association of Biological Standardization, 1987
40. Treasurer and Member of Executive Committee, International Association of Gerontology,
1985 - 1989.
41. Member, Board of Directors, The Center for Climacteric Studies, Inc., Gainesville, FL,
1985 - 1988.
42. Treasurer and Member of Executive Committee, International Association of Gerontology,
1985 - 1989.
43. Senior Editor, Biological Sciences, The Microfiche Collection of Information on
Gerontology and Geriatric Medicine, University Microfilms International, Ann Arbor,
Michigan, 1984 - 1990.
44. Gerontological Society of America, Committee Member for Glenn Foundation Award in
Basic Biological Research in Aging, 1990.
45. Referee, Macy Faculty Scholar award Program, Josiah Macy Jr. Foundation, 1977 - 1987.
46. Gerontological Society of America, Committee Member for Development and Fund Raising,
1990 - 1991.
47. Chairman, Cell Culture Committee of the International Association of Microbiological
Standardization, Geneva, Switzerland, 1985 - 1993.
48. Member, Editorial Board, "Proceedings of the Society for Experimental Biology and
Medicine" 1987 - 1994.
49. Member, Advisory Committee, Center for Aging Research, University of California, Santa
Barbara, CA, 1981 - 1990.
50. Member, Scientific Advisory Board, American Longevity Association, Inc., University of
California, Los Angeles, CA, 1981 - 1990.
51. Vice President, Member of Board of Directors and Executive Committee, Chairman of the
Research Advisory Committee (Study Section), American Federation for Aging Research, New
1981 - 1995.
52. Member, Board of Directors and Editorial Board, Bollettino Dell "Instituto
Sieroterapico Milanese, Archivo de Microbiologia de Immunologia," Milan, Italy, 1968
53. California Foundation for Medical Research, Member, 1979 - 1985.
54. Member, Editorial Board, "A Revista Portuguesa de Medicina Geriatrica", 1987
55. Member, Advisory Board, Lousiana Gerontology Education Center, Louisiana State
University, New Orleans, LA, 1989 - 1994.
56. Gerontological Society of America, Chairman, Working Group to Establish Executive
Director Job Description and Search Committee for Executive Director, 1991-1992.
57. Chairman, Search Committee, For Executive Director of the Gerontological Society of
58. Visiting Professor at Oita Medical University, Oita, Japan, 1993 - 1995.
Scientific Journal Affiliations
1. Editor-in-Chief, "Experimental Gerontology", Published by Elsevier Press,
N.Y., 1984 - 1998.
2. Member, Editorial Board, "Mechanisms of Ageing and Development", 1972
3. Foreign Reviewing Editor, "Tissue Culture Research Communications" (Journal
of the Japanese Tissue Culture Association) 1991 - Present.
4. Member, Editorial Board, "Proceedings of the Society for Experimental Biology and
Medicine," 1988 - 1992.
5. Member, Board of Directors and Editorial Board, "Bollettino Dell Instituto
Sieroterapico Milanese, Archivo de Microbiologia ed Immunologia'", Milan, Italy, 1968
6. Member, Editorial Board, "Executive Health Report", 1970 -Present.
7. Member, Editorial Board, "A Revista Portuguesa de Medicina Geriatrica", 1987
8. Member, Editorial Board, "Journal of Bacteriology", 1964-1972.
9. Member, Editorial Board, "Journal of Virology", 1967-1970.
10. Member, Editorial Board, "Infection and Immunity" 1968-1978.
11. Assistant Editor, "In Vitro", Journal of the Tissue Culture Association,
13. Editor, Biological Sciences Section, "Journal of Gerontology", 1975-1980.
14. Member, Editorial Advisory Board, "Protocols in Cell and Tissue Culture,"
John Wiley and Sons, West Sussex, England, 1989 - Present.
15. Member, Editorial Board, Cell Structure and Function (Journal of the Japan Society for
16. Member, Advisory Board, Geriatrics and Gerontology International, Tokyo, Japan,
Blackwell Publishing Co.
17. Honorary Editor, Experimental Gerontology, 2000 to Present
1. Professor of Anatomy, University of California School of Medicine, San Francisco,
California. 1988 - Present.
2. Professor of Zoology and of Microbiology and Immunology, Center for Gerontological
Studies, School of Liberal Arts and Sciences and School of Medicine, University of
Florida, Gainesville, Florida, 1981 - 1986.
3. Professor of Medical Microbiology, Stanford, University School of Medicine, Stanford,
California, July, 1968 - February, 1976.
4. Sponsor and Instructor of National Institute of Child Health and Human Development
Summer Courses on the Biology of Aging, 1970; Instructor, 1969 - 1973.
5. Assistant Professor, Department of Research Medicine, School of Medicine, University of
6. Organizer and Instructor in Course in Principles and Applications of Cell Culture,
Department of Medical Microbiology, University of the West Indies, Kingston, Jamaica, West
7. Assistant Director of the Tissue Culture Association Summer Course, University of
8. Organizer and Instructor in Tissue Culture Course in Graduate School, School of
Medicine, University of Puerto Rico, San Juan, Puerto Rico, 1962-1967.
9. Associate in Tissue Culture Association Summer Course, Madison, Wisconsin, 1961, on
"Principles and Application of Tissue Culture."
10. Organizer and Associate in Medical Microbiology in course "Principles and
Application of Tissue Culture" in Graduate School of Arts and Sciences, University of
Pennsylvania, Department of Medical Microbiology, 1959-1960.
11. Assistant Instructor in Medical Microbiology, University of Pennsylvania, June 1955 -
12. Visiting Professor, Kurume University Medical School, 1991- Present.
13. Visiting Professor, Oita Medical University, Oita, Japan 1991-1995.
14. National Science Foundation Chautauqua Course Organizer and Lecturer, 1998, 1999,
2000, 2002. How and Why We Age, Temple University, Philadelphia, PA
Honors and Awards
1. Career Development Awardee of the National Cancer Institute, National Institutes of
Health, Bethesda, Maryland, 1962-1970.
3. Fifth Pomerat Memorial Lecturer on Cultured Human Cells and the Biology of Senescence,
University of Texas Medical Branch, Galveston, Texas, February, 1970.
4. Fourteenth Annual Ciba Foundation Lecturer on Research in Aging, Leeds, England, April,
5. Commission for Aging Research Lecturer on Cell Aging and Cell Differentiation. The
Academy of Sciences and Literature, Mainz, Germany, November, 1971.
6. Huddleson Memorial Lectureship in Microbiology, Michigan State University, 1967.
7. Recipient of annual Robert Kleemeier Award, Gerontological Society of America, 1972,
for basic research in the biology of aging.
8. University of Southern California, Gerontology Center, Biomedical Sciences and Aging
9. University of Southern California, Gerontology Center, Kesten Lecture Award, 1974,
"for pioneering findings in microbiology that will open the door for more research
avenues on the problem of aging."
10. The Albert H. Rowe, Sr., M.D. Lecture, Samuel Merritt Hospital, Thirtieth Annual
Medical Seminar, Oakland, California, 1975.
11. Non-Resident Fellow, Institute for Higher Studies, Santa Barbara, CA
12. Visiting Scientist, Weizmann Institute of Science, Center for Aging, Rehovoth, Israel,
July 1975 and September 1980.
13. Keynote speaker, Centennial Celebration of the Founding of the Free University of
Amsterdam and Thirtieth Anniversary of the Founding of the School of Medicine, Amsterdam,
The Netherlands, September, 1980.
15. Recipient of $20,000 Brookdale Award for Biological and Clinical Research from the
Gerontological Society of America, November 1980, "for nationally and internationally
recognized scholarly and scientific contributions to biological and clinical research in
16. American Federation for Aging Research, Leadership award, New York City, May 1983,
"In recognition of his pioneering research in the fields of geriatrics and
gerontology and, in particular, his seminal studies on the life cycles of cell growth and
reproduction. His work has profoundly influenced the direction of present and future
investigations into the biomedical mechanisms of the aging process."
17. Elected President, Gerontological Society of America, 1982-1983.
18. Academy of Science and Literature, Mainz, Germany, and Johannes Gutenberg University,
Mainz, Thirtieth Karl August Forster Honorary Lectureship in Programmed Biosynthesis,
Mainz, Germany, May, 1983.
19. The 1984 Hoffman - LaRoche Lectureship in Microbiology, Waksman Institute of
Microbiology, Rutgers Univ., Rutgers, N.J., April, 1984.
20. The 1984 Wadsworth Memorial Fund Lecturer, Rush Medical College, Rush-Presbyterian-St.
Luke's Medical Center, Chicago, Illinois, April, 1984.
21. The 1984 Presidential Citation Award of the International Organization for
Mycoplasmology for "Discovery and Identification of the Etiological Agent of Human
Primary Atypical Pneumonia, Mycoplasma pneumoniae", Jerusalem, Israel, June, 1984.
22. The 1984 Samuel Roberts Noble Foundation Research Recognition Award, Ardmore,
Oklahoma, October, 1984. Prize of $5000 and Silver Bowl for "discovery of the limited
proliferative capacity of cultured normal human cells, establishment of the field of
cytogerontology, establishment of the WI-38 cell strain used
world-wide for human virus vaccine products, and for discovery and identification of the
etiological agent of human primary atypical pneumonia, Mycoplasma pneumoniae."
23. Listed in "Who's Who in America" and "International Who's Who in
Medicine", "American Men and Women of Science", etc.
24. Elected Councilor of the Society for Experimental Biology and Medicine, 1984-1988.
25. The Tenth Stuart Mudd Memorial Lecture Award, Sponsored by the Eastern Pennsylvania
Chapter of the American Society of Microbiology, 29 April 1985, for "discovery of
Mycoplasma pneumoniae as the etiologic agent of primary atypical pneumonia, development of
the first human diploid cell strain, and the
"Hayflick Limit" that occurs in the limited ability of cultured normal human and
animal cells to divide."
26. Invited speaker, 1986 and 1992 Sandoz Lectures in Gerontology, Basel, Switzerland,
February, 1986 and February 1992.
27. June 1986, Elected Fellow of the American Association for the Advancement of Science
"for fundamental studies on the finite lifetime of cultured normal human cells
interpreted as cell aging and for discovery of a new mycoplasma causing human
28. One of the 1000 most cited contemporary scientists in the world in the fields of
biochemistry, biophysics, cell biology, enzymology, genetics and molecular biology,
Institute for Scientific Information, Current Contents, #41, October 12, 1981, page 5.
29. Author of 4 of the 100 most cited scientific papers of the 2 million papers published
in the basic biomedical sciences from 1961 to 1978, Institute for Scientific Information,
Current Contents, #5, February 4, 1980, page 5.
30. Rosenfield Program in Public Affairs, International Relations and Human Relations,
Honorary Lecturer, Grinnell College, Grinnell, Iowa, April, 1989.
31. Elected to Honorary Membership in the Tissue Culture Association by the Council,
Executive Board and Membership, April, 1989. Honorary Membership is the highest
award that the Association can confer upon a person in recognition of their outstanding
contributions to the field of cell and tissue culture. Only 23 individuals have been so
honored." (From notice of award.)
32. Chairman and Speaker, Italian Society of Gerontology and Geriatrics, Regional Congress
on "Longevity," Universita "La Sapienza," Rome, Italy, October 1990
33. Invited Speaker on "Cellular Aging" at Istituto Scientifico Roussel, Italia,
Colloquium on Aging, Rome, Italy, Villa Cedro, October 1990.
34. Co-winner (with Prof. Hans Thomae, Bonn, Germany) of the 1991 International
Association of Gerontology Sandoz Prize in Gerontology. (50,000 Swiss Franks)
35. Bernard M. Bloomfield Memorial Lecturer on the occasion of the dedication of the
Bloomfield Centre for Research on Aging, Lady Davis Institute for Medical Research, The
Sir Mortimer B. Davis Jewish General Hospital, Montreal, Quebec, Canada, October 2, 1991.
36. Lectura Magistralis, "The Cell Biology of Human Aging," 20th National
Congress, Italian Society of Pathology, Modena, Italy,October 1990.
37. Visiting Professor (Professore Contratto) University of Parma, Faculty of Medicine amd
Surgery, Parma, Italy, October, 1991.
38. Elected Honorary Member, Society of Medicine and Natural Science of the University of
Parma, Parma, Italy, October 1991. Medal Awarded.
39. Honorary Lecture, "Biology of Aging," Ateneo di Brescia Accademia di Scienze
Lettere ed Arti and the Centro di Studio e Ricerca Sulla Terza Eta', Universita degli
Studi di Brescia, Italy, October 1991.
40. Honorary Lecture, "Cell Biology of Aging," University of Pisa, Pisa, Italy,
41. Recipient of Presidential Award Silver Medal from the International Association of
Gerontology (IAG) for "distinguished contributions as Treasurer of the IAG,
42. Visiting Professor, Kurume University Medical School, 1991-Present
43. Visiting Professor, Oita Medical University, Oita, Japan 1991-1995.
44. Eighth Annual Walter A. Rohlfing Medical Lectureship, March 18, 1995, University of
California, Fresno, Medical Education Program
45. Benjamin Leiberman Memorial Lecture, October 16, 1995, University of California, San
Francisco-Mount Zion Center on Aging.
46. Distinguished Lifetime Achievement Award of the Society for In Vitro Biology for
your exemplary record of achievement and contribution in the pioneering work on how
mammalian cells age, which has been instrumental in gerontology and medical science.
Awarded at the World Conference on Cell Culture, San Francisco, CA, June, 1996.
47. Elected Academician by the Ukrainian Acadamy of Medical Sciences, Kiev, Ukraine,
48. Annual Hayflick Lectureship established by the American Aging Association,
49. Recipient of the Herbert De Vries Research Award of the Council on Aging
and Adult Development of the American Alliance for Health and Physical Education In
recognition of outstanding research and writing that has contributed to the Council on
Aging and Adult Development and to the Profession. Presented at Annual Meeting, St.
Louis, MO, March 1997.
50. Sigma Xi, Honorary Scientific Society, Endowed Lecturship, University of Southern
Illinois, Carbondale, Illinois, October 16, 1997.
51. Elected corresponding member of the Société de Biologie of France, 1998. The
Society, was founded in 1864 with headquarters at the Collége de France, Paris.
53. Keynote Speaker, Tenth Biennial Conference of the Hawaii-Pacific Gerontological
Society, Honolulu, Hawaii, September, 1998.
54. Keynote Speaker, International Conference on Telomeres and Telomerase, Marianne
Leibert, Inc., Redwood City, CA, June,1998.
55. Keynote Speaker, Fourth International Conference on Neurobiology and
Neuroendocrinology of Aging, Bregenz, Austria, July, 1998.
56. Dr. Toon van Wezel Award and Honorary Lecture, for Contributions made over
several decades to the science and practical application of mammalian cell cultures.
Awarded by The European Society for Animal Cell Technology (ESACT), Lugano, Switzerland,
25-29 April 1999.
57. Lord Cohen of Birkenhead Medal presented by the British Society for Research on Aging
for major contributions to Gerontology, London, June, 1999.
58. Keynote Lecture, FDA Conference on Evolving Scientific and Regulatory Perspectives on
Cell Substrates for Vaccine Development, Rockville, MD, September 7, 1999.
59. Boureston Lecturer, Keynote Address, Minnesota Gerontological Society Annual Meeting,
Minneapolis, April 28, 2000.
60. Keynote Speaker, California Society of Plastic Surgeons, Santa Monica, CA, May 26,
61. Annual Hayflick Lecture in Aging established at the University of Alabama at
Birmingham and presentation of commemorative plaque, October 2000.
62. Publication of Hayflick, his limit, and cellular ageing by J. Shay and W.
Wright, a Timeline, in Nature Reviews, Molecular Cell Biology, October 1, 2000, pages
63. Recipient of the year 2001, $10,000 Life Extension Prize and Laureate Diploma from the
Regenerative Medicine Secretariat for " ..your discovery of the finite replicative
capacity of normal human diploid cells and his contribution to defining the relation of
replicative potential to organismal aging." and "..not only has the importance
of your discovery been grossly ignored by our scientific-medical culture but also that the
significance of your work needs to be duly recognized."
64. 2003 Joe L. Parkin Honorary Memorial Lecturer, University of Iowa College of Medicine,
Iowa City, Iowa
65. Invited Speaker at Nobel Conference XL, The Science of Aging, Gustavus
Adolphus College, Oct.5-6, 2004,St.Peter MN
66. Invited to officially open and dedicate the Henry Wellcome Laboratory for
Biogerontology Research at The University of Newcastle upon Tyne, November 10, 2004.
1. Senate Committee on Government Operations, Senator Abraham Ribicoff, Chairman, 1973,
2. Subcommittee on Federal, State and Community Services on the Select Committee on Aging,
House of Representatives, 95th Congress, February, 1978, Washington, D.C., Representative
Claude Pepper, Chairman.
3. Subcommittee on Labor, Health and Human Services and Education of the Committee on
Appropriations, House of Representatives, Representative Natcher, Chairman, May 1983.
4. Subcommittee on Pensions of the Senate Committee on Finance, Senator Chaffee, Chairman,
Memberships in Scientific Societies
American Society for Microbiology (Emeritus)
American Association for the Advancement of Science (Fellow)
Tissue Culture Association (Vice President, 1970) (Emeritus) (Renamed Society for In Vitro
American Society for Cell Biology (Emeritus)
Gerontological Society of America (President 1982-83)
Society for Experimental Biology and Medicine (Councilor, 1984-1988, Editorial Board
Member, 1988- 1994) (Emeritus)
Association for the Advancement of Aging Research (Fellow)
American Aging Association
American Cancer Society (Emeritus)
International Association of Microbiological Standardization (Chairman, Cell Culture
International Organization for Mycoplasmology (Retired)
American Society on Aging (Board of Directors)
American Federation for Aging Research (Board of Directors, Former Chairman, Scientific
Advisory Board and Member, Executive Committee)
Federation of American Societies for Experimental Biology (FASEB)
Tissue Culture Association, California Chapter, (President 1970)
Aging Prevention Research Foundation (Scientific Advisory Board)
American Association for Cancer Research
American Association of Pathologists
California Foundation for Biomedical Research
American Longevity Association (Scientific Advisory Board)
European Society for Animal Cell Technology (ESACT) (Retired)